Prof. Edward A. Codling


Professor of Mathematical Biology

Department of Mathematical Sciences

University of Essex, UK

Back to Homepage


Editorial and Reviewing


As well as producing my own publications, I review for 41 different journals in the fields of applied mathematics, ecology and environmental sciences:

ANZIAM Journal, Applied Mathematical Modelling, Aquatic Living Resources, American Naturalist, Animal Behaviour, Behavioral Ecology and Sociobiology, Bulletin of Mathematical Biology, Contributions to Zoology, Current Zoology, Ecological Applications, Ecological Complexity, Ecological Modelling, Ecology, Ecology Letters, Environmental Management, Estuaries and Coasts, Fish and Fisheries, Fisheries Research, Functional Ecology, Global Change Biology, ICES Journal of Marine Science, Journal of Animal Ecology, Journal of Biological Dynamics, Journal of Marine Systems, Journal of Mathematical Biology, Journal of Plankton Research, Journal of the Royal Society Interface, Journal of Spatial Information Science, Journal of Theoretical Biology, Mathematical Modelling of Natural Phenomena, Methods in Ecology and Evolution, Movement Ecology, Oikos, Physica A, PLoS Computational Biology, PLoS ONE, Proceedings of the National Academy of Sciences (PNAS), Proceedings of the Royal Society A, Proceedings of the Royal Society B, Theory in Biosciences, Trends in Ecology and Evolution.


I am a member of the editorial board of Ecological Complexity.

I have also acted as guest subject matter editor for Ecological Applications.

I am guest editor of two journal special issues:

     Interface Focus (April 2012)

     Ecological Complexity (Dec 2013)





∞  Submitted
51. Berdahl, AM, Kao, AB, Flack, A, Westley, PAH, Codling, EA, Couzin, ID, Dell, AI, Biro, D. Collective animal navigation and the emergence of migratory culture.
50. Bailey, JD, Wallis, J, Codling, EA. Navigational efficiency in a biased and correlated random walk model of individual animal movement.
49. Bode, NWF, Codling, EA. Exploring determinants of pre-movement delays in a virtual crowd evacuation experiment.
48. Barker, ZE, Vazquez Diosdado, JA, Bell, NJ, Hodges, HJ, Codling, EA, Croft, DB, Amory, JR. Use of a novel sensor combining local positioning and acceleration to measure feeding behaviour changes associated with lameness in dairy cattle.
47. Walker, ND, Susanto, H, Steinke, M, Codling, EA. Top-down and bottom-up control in a multitrophic system: the role of nutrient limitation and infochemicals in a plankton food web model.
  In Press


Piercy, JJB, Smith, DJ, Simpson, SD, Jompa, J, Codling, EA. High temporal resolution sampling reveals reef fish settlement is highly clustered. Canadian Journal of Fisheries and Aquatic Sciences, in press.
  Published Journal Articles


Auger-Mth, M, Derocher, AE, DeMars, C, Plank, MJ, Codling, EA, Lewis, MA. 2016. Evaluating random search strategies in three mammals from distinct feeding guilds. Journal of Animal Ecology, 85: 1411-1421.
44. Pazhayamadom, DG, Kelly, CJ, Rogan, E, Codling, EA. 2016. Self-Starting Cumulative Sum Harvest Control Rule (SS-CUSUM-HCR) for managing data limited fisheries. Canadian Journal of Fisheries and Aquatic Sciences, 73: 366-381.
43. Codling, EA, Bode, NWF. 2016. Balancing direct and indirect sources of navigational information in a leaderless model of collective animal movement. Journal of Theoretical Biology, 394: 32-42.
42. Bode, NWF, Miller, J, O'Gorman, R, Codling, EA. 2015. Increased costs reduce reciprocal helping behaviour of humans in a virtual evacuation experiment. Scientific Reports, 5: 15896. (pdf)
41. Auger-Mth, M, Derocher, AE, Plank, MJ, Codling, EA, Lewis, MA. 2015. Differentiating the Lvy walk from a composite correlated random walk. Methods in Ecology and Evolution, 6: 1179-1189. (pdf)
40. Kraak, SBM, Reid, DG, Bal, G, Barkai, A, Codling, EA, Kelly, CJ, Rogan, E. 2015. RTI outperforms traditional management in simulated mixed fisheries and cases incorporating protection of vulnerable species and areas. Fisheries Research, 172: 209-224. (pdf)
39. Pazhayamadom, DG, Kelly, CJ, Rogan, E, Codling, EA. 2015. Decision Interval Cumulative Sum Harvest Control Rules (DI-CUSUM-HCR) for managing fisheries with limited historical information. Fisheries Research, 171: 154-169. (pdf)
38. Vazquez Diosdado, JA, Barker, ZE, Hodges, HR, Amory, JR, Croft, DP, Bell, NJ, Codling, EA. 2015. Classification of behaviour in housed dairy cows using an accelerometer-based activity monitoring system. Animal Biotelemetry, 3: 15. (pdf)
37. Bode, NWF, Kemloh Wagoum, AU, Codling, EA. 2015. Information use by humans during dynamic route choice in virtual crowd evacuations. Royal Society Open Science, 2: 140410. (pdf)
36. Piercy, JJB, Codling, EA, Hill, AJ, Smith, DJ, Simpson, SD. 2014. Habitat quality affects sound production and likely distance of detection on coral reefs. Marine Ecology Progress Series, 516: 35-47. (pdf)
35. Codling, EA. 2014. Pest insect movement and dispersal as an example of applied movement ecology (Comment). Physics of Life Reviews, 11: 533-535. (pdf)
34. Codling, EA, Bode, NWF. 2014. Copycat dynamics in leaderless animal group navigation. Movement Ecology, 2: 11. (pdf)
33. Auger-Mth, M, Plank, MJ, Codling, EA. 2014. Distinguishing between Lvy walks and strong alternative models: comment. Ecology, 95: 1104-1109. (pdf)

Reply article by Reynolds

32. Kraak, SBM, Reid, DG, Codling, EA. 2014. Exploring the RTI (Real-Time Incentive) tariff-based approach to single-species fisheries management. Fisheries Research, 155: 90-102. (pdf)
31. Bode, NWF, Kemloh Wagoum, AU, Codling, EA. 2014. Human responses to multiple sources of directional information in virtual crowd evacuations. Journal of the Royal Society Interface, 11: 20130904. (pdf)
30. Lewis, ND, Morozov, A, Breckels, MN, Steinke, M, Codling, EA. 2013. Multitrophic interactions in the sea: assessing the effect of infochemical-mediated foraging in a 1-d spatial model. Mathematical Modelling of Natural Phenomena, 8: 25-44. (pdf)
29. Simpson, SD, Piercy, JJB, King, J, Codling, EA. 2013. Modelling larval behaviour and population dynamics of coral reef fishes. Ecological Complexity, 16: 68-76. (pdf)
28. Lewis, ND, Breckels, MN, Steinke, M, Codling, EA. 2013. Role of infochemical mediated zooplankton grazing in a phytoplankton competition model. Ecological Complexity, 16: 41-50. (pdf)
27. Codling, EA, Dumbrell, AJ. 2013. Bridging the Gap between Theory and Data in Ecological Models. Ecological Complexity, 16: 1-8. (pdf)
26. Bode, NWF, Codling, EA. 2013. Human exit route choice in virtual crowd evacuations. Animal Behaviour, 86: 347-358. (pdf)
25. Breckels, MN, Bode, NWF, Codling, EA, Steinke, M. 2013. The effect of grazing-mediated DMS production on the behavior of the copepod Calanus helgolandicus. Marine Drugs, 11: 2486-2500. (pdf)
24. Pazhayamadom, DG, Kelly, CJ, Rogan, E, Codling, EA. 2013. Self starting CUSUM approach for monitoring data poor fisheries. Fisheries Research, 145: 114-127. (pdf)
23. Lewis, ND, Breckels, MN, Archer, SD, Morozov, A, Pitchford, JW, Steinke, M, Codling, EA. 2012. Grazing-induced production of DMS can stabilize food-web dynamics and promote the formation of phytoplankton blooms in a multitrophic plankton model. Biogeochemistry, 110: 303-313. (pdf)
22. Knell, AS, Codling EA. 2012. Classifying area-restricted search (ARS) using a partial sum approach. Theoretical Ecology, 5: 325-339. (pdf)

Supplementary material

21. Bode, NWF, Franks, DW, Wood, AJ, Piercy, JJB, Croft, DP, Codling, EA. 2012. Distinguishing social from non-social navigation in moving animal groups. American Naturalist, 179: 621-632. (pdf)

Supplementary material and data

20. Kraak, SBM, Reid, DG, Gerritsen, HD, Kelly, CJ, Fitzpatrick, M, Codling, EA, Rogan, E. 2012. 21st century fisheries management: A spatiotemporally explicit tariff-based approach combining multiple drivers and incentivising responsible fishing. ICES Journal of Marine Science, 69: 590-601. (pdf)
19. Codling, EA, Dumbrell, AJ. 2012. Mathematical and Theoretical Ecology: linking models with ecological processes. Interface Focus, 2: 144-149. (pdf)
18. Plank, MJ, Codling EA. 2011. Sampling rate and misidentification of Lvy and non-Lvy movement paths: reply. Ecology, 92: 1701-1702. (pdf)
17. Codling, EA, Plank, MJ. 2011. Turn designation, sampling rate and the misidentification of power-laws in movement path data using maximum likelihood estimates. Theoretical Ecology, 4: 397-406. (pdf)

Supplementary material

16. Boakes, DE, Codling, EA, Thorn, GJ, Steinke, M. 2011. Analysis and modelling of swimming behaviour in Oxyrrhis marina. Journal of Plankton Research, 33: 641-649. (pdf)
15. Codling, EA, Bearon, RN, Thorn, GJ. 2010. Diffusion about the mean drift location in a biased random walk. Ecology, 91: 3106-3113. (pdf)  

Ecological Archives E091-215-A1 (Online Appendix) 

14. Kraak, SBM, Kelly, CJ, Codling, EA, Rogan, E. 2010. On scientists’ discomfort in fisheries advisory science: the example of simulation-based fisheries Management Strategy Evaluations. Fish and Fisheries, 11: 119-132. (pdf)
13. Breckels, MN, Boakes, DE, Codling, EA, Malin, G,  Archer, SD, Steinke, M. 2010. Modelling the concentration of exuded dimethylsulphoniopropionate (DMSP) in the boundary layer surrounding phytoplankton cells. Journal of Plankton Research, 32: 253-257. (pdf)

Reply article by Strom & Fredrickson

12. Plank, MJ, Codling, EA. 2009. Sampling rate and misidentification of Lvy and non-Lvy movement paths. Ecology, 90: 3546-3553. (pdf)  

Ecological Archives E090-248-A1 (Online Appendix)  -  Ecological Archives E090-248-A2 (Online Appendix)

Ecological Archives E090-248-A3 (Online Appendix)

11. Faria, JJ, Codling, EA, Dyer, JRG, Trillmich, F, Krause, J. 2009. Navigation in human crowds: testing the many-wrongs principle. Animal Behaviour, 78: 587-591. (pdf)      

Animal Behaviour Featured Articles - September 2009

10. Le Quesne, WJF, Codling, EA. 2009. Managing mobile species with MPAs: the effects of mobility, larval dispersal, and fishing mortality on closure size. ICES Journal of Marine Science, 66: 122-131. (pdf)
9. Codling, EA, Plank, MJ, Benhamou, S. 2008. Random walk models in biology. Journal of the Royal Society Interface, 5: 813–834. (pdf)

Faculty of 1000 Prime Recommended         Access the recommendation on F1000Prime               

8. Codling, EA. 2008. Individual-based movement behaviour in a simple marine reserve – fishery system: why predictive models should be handled with care. Hydrobiologia, 606: 55-61. (pdf)
7. Pitchford, JW, Codling, EA, Psarra, D. 2007. Uncertainty and sustainability in fisheries and the benefits of marine protected areas. Ecological Modelling, 207: 286-292. (pdf)
6. Codling, EA, Pitchford, JW, Simpson, SD. 2007. Group navigation and the 'many wrongs principle' in models of animal movement. Ecology, 88: 1864-1870. (pdf)   

Ecological Archives E088-111-A1 (Online Appendix)

5. Kelly, CJ, Codling, EA. 2006. ‘Cheap and Dirty’ Fisheries Science and Management in the North Atlantic. Fisheries Research, 79: 233-238. (pdf)
4. Kelly, CJ, Codling, EA, Rogan, E. 2006. The Irish Sea cod recovery plan: some lessons learned. ICES Journal of Marine Science, 63: 600-610. (pdf)
3. Codling, EA, Hill, NA. 2005. Calculating spatial statistics for velocity jump processes with experimentally observed reorientation parameters. Journal of Mathematical Biology, 51: 527-556. (pdf)
2. Codling, EA, Hill, NA. 2005. Sampling rate effects on measurements of correlated and biased random walks. Journal of Theoretical Biology, 233: 573-588. (pdf)
1. Codling, EA, Hill, NA, Pitchford, JW, Simpson, SD. 2004. Random walk models for the movement and recruitment of reef fish larvae. Marine Ecology Progress Series, 279: 215-224. (pdf)
∞  Published Conference Proceedings & Abstracts
2. Bode, NWF, Codling, EA. 2016. Statistical models for pedestrian behaviour in front of bottlenecks. Traffic and Granular Flow '15 (pp. 81-88), Springer.
1. Barker, ZE, Hodges, HR, Vazquez Diosdado, JA, Bell, NJ, Codling, EA, Croft, DP, Amory, JR. 2016. Detection of dairy cow behaviour changes associated with claw horn lesions using biosensors. Advances in Animal Biosciences, 7:1.
∞  Book Chapters
2. Piercy, JJB, Smith DJ, Codling, EA, Hill, AJ, Simpson, SD. 2016. The Good, the Bad, and the Distant: Soundscape Cues for Larval Fish. In A.N.Popper & A. Hawkins (eds.) The Effects of Noise on Aquatic Life II (pp. 829-837), Springer, New York.
1. Plank, MJ, Auger-Mth, M, Codling, EA. 2013. Lvy or not? Analysing positional data from animal movement paths. In M. Lewis, P.K. Maini, & S.V. Petrovskii (eds.) Dispersal, Individual Movement and Spatial Ecology: a Mathematical Perspective (pp. 33-52), Springer.
1. Codling, EA, Kelly, CJ (eds). 2006. Reports from the FSS mini-symposia 2004-2005. Galway: Marine Institute. ISBN 1-902895-29-0 (pdf)
∞  Other Publications
5. Codling, EA, Thorn, GJ. 2009. Simplifying assessment and management of fisheries. Fishing News (31/07/2009), p10. (pdf)
4. Codling, EA, Kelly, CJ. 2006. F-PRESS: a stochastic simulation tool for developing fisheries management advice and evaluating management strategies. Irish Fisheries Investigation Series, No. 17 (34pp). (pdf)
3. Codling, EA, Kelly, CJ, Clarke, M. 2005. Comparison of the effects of exploitation on theoretical long-lived fish species with different life-history strategies and the implications for management. ICES CM 2005/N. (pdf)
2. Codling, EA, Kelly, CJ. 2005. Uncertainty in fisheries science and management. Fishing News (03/06/2005), p14. (pdf)
1. Codling, EA. 2003. Biased random walks in biology. PhD thesis, University of Leeds. (pdf - 5MB)



Page last updated September 2017